Custodians do it. Which is ironic because the definition of a custodian is: "a person who has responsibility for taking care of or protecting something". Any holder with "nominees" in the name is a 'custodian'. And guess what, the Top 5 are all custodians:
They would be your cuplrits, in my humble opinion. I used to work for the one of the custodians listed above (in IT, never on the business side). They get paid a fee to loan out stock for corporate activities, one of which is shorting. I've always thought there was a possible conflict of interest there. On the one hand they hold stock for shareholders trying to maximise value, on the other they loan it out for shorting.
ORE Price at posting:
$4.30 Sentiment: Buy Disclosure: Held